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A study found that a 50 year old TMV that had been in storage at 4 C could still infect tobacco plants.[5] Once the TMV has used the host ribosomes to synthesize the replicase-associated proteins, they attach themselves to the 3 end of the positive sense strand of the RNA and begin the synthesis of the negative sense strand of RNA.[5] The negative sense RNA acts as the template strand for the synthesis of a full positive sense genome and subgenomic RNA (sgRNA)