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Notably, berberine loaded in chitosan/SLN nanoparticles demonstrated a much slower cumulative release than free-berberine, suggesting the effective protective effect of nanoparticles on berberine against the gastrointestinal harsh acidic environment

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Both calpain and CDK5 contribute to GSK-3 activation: calpain by cleaving its inhibitors and CDK5 through direct phosphorylation, which phosphorylates DRP1, promoting mitochondrial fission [37, 48, 49]