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10.1242/dev.121.8.2265 Summary Keywords hedgehog, morphogen, Drosophila , wing disc, heparan sulfate, QCM-D, heparin, development Citation Manikowski D, Steffes G, Froese J, Exner S, Ehring K, Gude F, Di Iorio D, Wegner SV and Grobe K (2023) Drosophila hedgehog signaling range and robustness depend on direct and sustained heparan sulfate interactions