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Using unsupervised clustering and canonical markers [13], we identified basal cell populations highly expressing KRT17 and KRT5 (basal1 and basal2 clusters), club cell populations highly expressing MUC5B and SCGB1A1 (club cell and secretory club cell clusters), goblet cell populations highly expressing MUC5AC and TSPAN8 genes (goblet1,2,3 clusters) and ciliated cell populations highly expressing FOXJ1 and TUBB4B (ciliated 1, 2, 3, 4, 5 clusters) (Fig